Gravitropism Gets Germinating Seeds Oriented

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Toward the light. Away from gravity.

Germinating seedlings know which way is up. Perhaps they just know which way is down. They can not see, hear, taste, smell or feel anything like we can. Nonetheless, they know which direction to extend their first root and stem. One thing that they can perceive is gravity. Gravitropism, which was formerly known as geotropism, is how they respond to gravity, or the Earth that generates it.

Positive gravitropism is why the first root to emerge from a seed extends downward toward gravity. Negative gravitropism is why the first stem to emerge from a seed extends upward away from gravity. Positive is toward. Negative is away. Roots and stems that develop after the first, disperse in other directions in response to other stimuli, but never really forget where gravity comes from.

Once a primary stem of a seedling emerges from the soil, it immediately responds to sunlight. Just as it exhibits negative gravitropism to grow away from gravity, it exhibits positive phototropism to grow toward sunlight. Since sunlight comes from above, positive phototropism is compliant to negative gravitropism. Branches will later disperse laterally to avoid the shade of other branches.

While branches are finding their way in the World, roots are doing the same. All of them can not always reach for the center of the Earth. They branch and disperse laterally as they sense that they are sufficiently deep in the soil. Those that venture too deeply sense an inhibition of gas exchange. Those that are too shallow sense if they get too warm or dry. There are a few types of tropisms.

Roots respond to moisture, nutrients, chemicals, temperature and mechanical stimulation within the soil. Branches and foliage respond to humidity, wind, temperature variations and air pollution. In order to function within their dynamic situations, plants somehow coordinate their responses to all of the many stimuli they experience. They are impressively perceptive, as well as responsive.

Even celery, green onions and leafy tops of carrot in a refrigerator can reach upward away from gravity.

Tropism Gives Plants Some Direction

80523thumbWhen a seed germinates, the roots know that they want to go down, and the new stem knows that it wants to go up. Going up sounds simple enough. Stems just go towards the light. How do they know where the light is while they are still under the soil? Well, new stems go upward for the same but opposite reason that roots go downward; gravity. Roots go toward it. Stems go away from it.

Tropism is how plant parts respond to a variety of stimuli, particularly gravity and light, but also water, chemicals, mechanical stimulation, trauma and electricity. Response to gravity is geotropism. Roots exhibit positive geotropism by growing towards gravity. Stems exhibit negative geotropism by growing away from gravity. Stems exhibit positive phototropism by growing towards sunlight.

Roots are always figuring out where to go next by prioritizing their innate positive geotropism, their tropism for or against certain chemicals, and their tropism for moisture but against saturation. Until we see them migrating into lawns, displacing concrete or getting into a septic systems, their work underground is unseen. Tropisms above ground are quite visible and perhaps informative.

When a tall herbaceous plant falls over, it tries to get up. If unable to, it can at least curve new grow upward. Even cut flowers and vegetables in the kitchen can do that much. Snapdragons that are initially arranged leaning outwardly from the center of a floral arrangement can go vertical within a day or two. The fluffiest houseplants regularly get turned so than they do not favor one sunny side.

Trees are too big to move, but on rare occasion, it happens. The crossed pairs of Mexican fan palms commonly planted outside of In-N-Out Burger restaurants get planted at a lean while mature, but then grow vertically after installation, leaving an angular kink where the direction changed. Such a kink in an otherwise straight trunk of a tree that was not planted at an angle might indicate a sudden destabilization. A curved trunk indicates either a slow destabilization, or tropism to escape shade.